{"id":2471,"date":"2021-08-31T17:02:52","date_gmt":"2021-08-31T17:02:52","guid":{"rendered":"https:\/\/cjai.instances.ca\/?p=2471"},"modified":"2021-08-31T17:09:47","modified_gmt":"2021-08-31T17:09:47","slug":"ips-calligraphus","status":"publish","type":"post","link":"https:\/\/cjai.biologicalsurvey.ca\/fr\/ips-calligraphus\/","title":{"rendered":"Ips calligraphus"},"content":{"rendered":"
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Ips bark beetles of the World ><\/a><\/h4>\t\t<\/div>\n\t\t\t\t<\/div>\n\t\t\t\t
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Checklist><\/a><\/h4>\t\t<\/div>\n\t\t\t\t<\/div>\n\t\t\t\t
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Scientific Name<\/h1>

Ips calligraphus (Germar, 1824)<\/p>

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Synonyms<\/h1>

Bostrichus exesus<\/em>\u00a0Say, 1826<\/p>

Bostrichus chloroticus<\/i>\u00a0Dejean, 1837<\/p>

Bostrichus conformis<\/i>\u00a0Dejean, 1837<\/p>

Tomicus praemorsus<\/em>\u00a0Eichhoff, 1868<\/p>

Tomicus interstitialis<\/em>\u00a0Eichhoff, 1869<\/p>

Ips ponderosae<\/em>\u00a0Swaine, 1925<\/p>

Common names: Coarse writing engraver, six-spined Ips, six-spined engraver beetle (English)<\/p>

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Diagnostic notes<\/h1>

-Has\u00a0six spines on the elytral\u00a0declivity<\/a>\u00a0and\u00a0is similar to\u00a0I. apache<\/a><\/i>\u00a0Distinguished from other\u00a0Ips\u00a0<\/i>spp. by the presence of three spines after the third\u00a0declivity<\/a>\u00a0spine<\/a>.
-Potentially sympatric in Arizona and northern Mexico with related and\u00a0morphologically similar species\u00a0I. apache<\/a><\/i>.
–I. calligraphus\u00a0<\/em>is often larger (pronotal width 1.3\u00a0to 2.5 mm) than\u00a0I.\u00a0<\/em>apache<\/em><\/a>\u00a0<\/em>(pronotal width 1.3\u00a0to\u00a02.1 mm) and striations on the pars stridens are\u00a0broader (mean values:\u00a0I. apache<\/a><\/em>\u00a0= 0.9 \u00b5m;\u00a0I. calligraphus<\/em>\u00a0= 1.0 \u00b5m) (Lanier\u00a0et al<\/i>. 1991).<\/p>

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Morphological Summary<\/h1>

females
Body.\u00a0<\/b>3.8-5.9(-6.7) mm long, 2.6-2.9 times longer than wide;\u00a0pronotum<\/a>\u00a01.2-1.3 times longer than wide.
Head.\u00a0<\/b>Epistomal<\/a>\u00a0margin with uniseriate row of tubercles absent or present with elongate mesal tubercle.\u00a0Frons<\/a>\u00a0outline convex in lateral view; vestiture fine (not hiding part of integument); surface sculpture near epistoma densely tuberculate-punctate<\/a>;\u00a0central carina<\/a>\u00a0present or absent; central tubercle present and single, separated from base of\u00a0epistomal<\/a>\u00a0setae by 2-3 tubercle diameters, without pair of circular tubercles on either side of midline;\u00a0transverse carina<\/a>\u00a0absent;\u00a0frons<\/a>\u00a0central fovea present or absent; circular tubercles above top of eyes present – up to one third of all tubercles. Vertex and\u00a0pronotum<\/a>\u00a0with stridulatory apparatus (pars stridens).\u00a0Antennal club sutures<\/a>\u00a0acutely\u00a0angulate<\/a>.
Prothorax.\u00a0<\/b>Protibiae with four or five\u00a0socketed teeth<\/a>\u00a0on apical half (does not include apical\u00a0spine<\/a>).
Elytra.<\/strong>\u00a0Interstriae<\/a>\u00a0punctate<\/a>\u00a0(observed on\u00a0interstriae<\/a>\u00a02 and 3 on middle third of elytral\u00a0disc<\/a>), punctures 0.3(-0.4) times diameter of adjacent\u00a0strial punctures<\/a>\u00a0(punctures and\u00a0striae<\/a>\u00a0measured at steepest part of puncture wall), interstrial setae longer than width of scutellar shield,\u00a0interstriae<\/a>\u00a02-3 times as wide as adjacent\u00a0striae<\/a>. Elytral\u00a0declivity<\/a>\u00a0with six spines per side,\u00a0spine<\/a>\u00a03, 4 or 5 largest;\u00a0spine<\/a>\u00a01 (largest on 2nd interstria) closer to\u00a0suture<\/a>\u00a0than\u00a0spine<\/a>\u00a02; spines 1 and 2 separated at base by distance greater than height of\u00a0spine<\/a>\u00a01;\u00a0spine<\/a>\u00a02 closer to\u00a0spine<\/a>\u00a03 than\u00a0spine<\/a>\u00a01;\u00a0spine<\/a>\u00a03\u00a0tapered<\/a>\u00a0or straight sided with\u00a0tapered<\/a>\u00a0apex, apex acute or right-angled, with apical half symmetrical or asymmetrical in lateral view; spines 2 and 3 not on shared tumescence, not in line with spines 1 and 4 (posterodorsal view);\u00a0spine<\/a>\u00a04 closer to\u00a0spine<\/a>\u00a03 than\u00a0spine<\/a>\u00a05; declivital integument shiny.<\/p>

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males
Body.\u00a0<\/b>3.8-5.9(-6.7) mm long, 2.6-2.9 times longer than wide;\u00a0pronotum<\/a>\u00a01.2-1.3 times longer than wide.
Head.\u00a0<\/b>Epistomal<\/a>\u00a0margin with uniseriate row of tubercles uninterrupted medially or margin with uniseriate row of tubercles with elongate mesal tubercle.\u00a0Frons<\/a>\u00a0outline convex in lateral view; vestiture\u00a0fine (not hiding part of integument); surface sculpture near epistoma densely tuberculate-punctate<\/a>\u00a0or with isolated tubercles;\u00a0central carina<\/a>\u00a0absent; central tubercle present and single, separated from base of\u00a0epistomal<\/a>\u00a0setae by 2-3 tubercle diameters, without pair of circular tubercles on either side of midline;\u00a0transverse carina<\/a>\u00a0absent;\u00a0frons<\/a>\u00a0central\u00a0fovea<\/a>\u00a0absent; circular tubercles above top of eyes present – up to one third of all tubercles. Vertex and\u00a0pronotum<\/a>\u00a0without\u00a0stridulatory apparatus<\/a>\u00a0(pars stridens).\u00a0Antennal club sutures<\/a>\u00a0acutely\u00a0angulate<\/a>.
Prothorax.\u00a0<\/b>Protibiae with three\u00a0socketed teeth<\/a>\u00a0on apical half (does not include apical\u00a0spine<\/a>).
Elytra.<\/strong>\u00a0Interstriae<\/a>\u00a0punctate<\/a>\u00a0(observed on\u00a0interstriae<\/a>\u00a02 and 3 on middle third of elytral\u00a0disc<\/a>), punctures 0.3(-0.4) times diameter of adjacent\u00a0strial punctures<\/a>\u00a0(punctures and\u00a0striae<\/a>\u00a0measured at steepest part of puncture wall), interstrial setae longer than width of scutellar shield,\u00a0interstriae<\/a>\u00a02-3 times as wide as adjacent\u00a0striae<\/a>. Elytral\u00a0declivity<\/a>\u00a0with six spines per side,\u00a0spine<\/a>\u00a03 largest;\u00a0spine<\/a>\u00a01 (largest on 2nd interstria) closer to\u00a0suture<\/a>\u00a0than\u00a0spine<\/a>\u00a02; spines 1 and 2 separated at base by distance greater than height of\u00a0spine<\/a>\u00a01;\u00a0spine<\/a>\u00a02 closer to\u00a0spine<\/a>\u00a03 than\u00a0spine<\/a>\u00a01;\u00a0spine<\/a>\u00a03 pedunculate (capitate) or\u00a0hooked<\/a>, apex right-angled or obtuse to rounded, with apical half asymmetrical in lateral view; spines 2 and 3 not on shared tumescence, not in line with spines 1 and 4 (posterodorsal view);\u00a0spine<\/a>\u00a04 closer to\u00a0spine<\/a>\u00a03 than\u00a0spine<\/a>\u00a05; declivital integument shiny.<\/p>

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Geographic Distribution<\/h1>

Bahama Islands; Canada (Ontario, Quebec); Dominican Republic; Mexico (Hidalgo, Jalisco, Nuevo Leon); Philippines (introduced); Jamaica; USA (Alabama, Arizona, Arkansas, California, Colorado, Delaware, Florida, Georgia, Indiana, Louisiana, Maryland, Massachusetts, Michigan, Minnesota, Mississippi, Missouri, Montana, Nebraska, New Hampshire, New Jersey, New Mexico, New York, North Carolina, Ohio, Oklahoma, Pennsylvania, South Carolina, South Dakota, Tennessee, Texas, Utah, Virginia, West Virginia, Wisconsin, Wyoming).<\/p>

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Hosts<\/h1>

Pinus<\/em>\u00a0spp.<\/p>

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Notes<\/h1>

Clade formed by (((I. apache<\/a><\/em>\u00a0+ I.\u00a0calligraphus<\/em>) +\u00a0I. lecontei<\/a><\/em>) +\u00a0I. cribricollis<\/a><\/em>), see Cognato and Sun (2007).<\/p>

\u00a0<\/p>

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References<\/h1>

Cognato, A.I.2015. Biology, systematics, and evolution of\u00a0Ips<\/i>.\u00a0In<\/i>\u00a0Bark beetles: biology and ecology of native and invasive species.\u00a0Edited by<\/i>\u00a0F.E. Vega and R.W. Hofstetter. Elsevier, San Diego, California. Pp. 351\u2013370.<\/i><\/p>

Cognato, A.I. and Sun, J.H. 2007. DNA based cladograms augment the discovery of a new\u00a0Ips<\/i>\u00a0species from China (Coleoptera: Curculionidae: Scolytinae).\u00a0Cladistics<\/i>,\u00a023<\/b>: 539\u2013551.<\/p>

EPPO. No date. Data Sheets on Quarantine Pests:\u00a0Ips calligraphus<\/em>. European and Mediterranean Plant Protection Organization.<\/p>

Wood, S.L. 1982. The bark and ambrosia beetles of North and Central America (Coleoptera: Scolytidae), a taxonomic monograph.\u00a0Great Basin Naturalist Memoirs<\/i>,\u00a06<\/b>: 1\u20131359.<\/p>

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Internet resources<\/h1>

https:\/\/www.barkbeetles.info\/regional_chklist_target_species.php?lookUp=1704<\/a><\/p><\/div>

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Ips bark beetles of the World > Checklist> Scientific Name Ips calligraphus (Germar, 1824) Synonyms Bostrichus exesus\u00a0Say, 1826 Bostrichus chloroticus\u00a0Dejean, 1837 Bostrichus conformis\u00a0Dejean, 1837 Tomicus praemorsus\u00a0Eichhoff, 1868 Tomicus interstitialis\u00a0Eichhoff, 1869 Ips ponderosae\u00a0Swaine, 1925 Common names: Coarse writing engraver, six-spined Ips, six-spined engraver beetle (English) Diagnostic notes -Has\u00a0six spines on the elytral\u00a0declivity\u00a0and\u00a0is similar to\u00a0I. apache\u00a0Distinguished from […]<\/p>","protected":false},"author":1,"featured_media":0,"comment_status":"closed","ping_status":"closed","sticky":false,"template":"","format":"standard","meta":{"footnotes":""},"categories":[1],"tags":[],"acf":[],"_links":{"self":[{"href":"https:\/\/cjai.biologicalsurvey.ca\/fr\/wp-json\/wp\/v2\/posts\/2471"}],"collection":[{"href":"https:\/\/cjai.biologicalsurvey.ca\/fr\/wp-json\/wp\/v2\/posts"}],"about":[{"href":"https:\/\/cjai.biologicalsurvey.ca\/fr\/wp-json\/wp\/v2\/types\/post"}],"author":[{"embeddable":true,"href":"https:\/\/cjai.biologicalsurvey.ca\/fr\/wp-json\/wp\/v2\/users\/1"}],"replies":[{"embeddable":true,"href":"https:\/\/cjai.biologicalsurvey.ca\/fr\/wp-json\/wp\/v2\/comments?post=2471"}],"version-history":[{"count":0,"href":"https:\/\/cjai.biologicalsurvey.ca\/fr\/wp-json\/wp\/v2\/posts\/2471\/revisions"}],"wp:attachment":[{"href":"https:\/\/cjai.biologicalsurvey.ca\/fr\/wp-json\/wp\/v2\/media?parent=2471"}],"wp:term":[{"taxonomy":"category","embeddable":true,"href":"https:\/\/cjai.biologicalsurvey.ca\/fr\/wp-json\/wp\/v2\/categories?post=2471"},{"taxonomy":"post_tag","embeddable":true,"href":"https:\/\/cjai.biologicalsurvey.ca\/fr\/wp-json\/wp\/v2\/tags?post=2471"}],"curies":[{"name":"wp","href":"https:\/\/api.w.org\/{rel}","templated":true}]}}