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Paratypes, 198, all males, all but a few with aedeagus examined:
UNITED STATES: AL: Dale Co., Ft. Rucker Mil. Res., 31.33 -85.71, barrier pitfall trap, 4-I to 16-IV-1999, R. Turnbow, 1 (Turnbow); Ft. Rucker Mil. Res., 31.33 -85.71, berlese leaf litter, 3-VII-2005, R. Turnbow, 1 (Turnbow); Lee Co., Auburn, 32.62 -85.48, sifting forest litter, 20-III-1972, E. J. Kiteley, 1 (CNC); Auburn, 32.62 -85.48, sifting forest litter, 25-III-1972, E. J. Kiteley, 1 (CNC). AR: Arkansas Co., White River N.W.R., 34.33 -91.12, woodland pitfall trap, 9-VII-1969, Underwood and Brown, 1 (UAAM); Ashley Co., pine forest, pitfall trap, Line 15 (A-15), 5-VIII-1974, Heiss, Howard and Chenowith, 1 (UAAM); 13.5 mi S of Crossett, 33.00 -91.83, pitfall trap, 74-2459 Line 13, 24-VI-1974, Heiss, Howard and Chenowith, 1 (UAAM); Bradley Co., 16-V-1946, 1 (UAAM); 19-VI-1963, 1 (UAAM); 5-VI-1965, 1 (UAAM); 24-VII-1965, 1 (UAAM); 30-VII-1965, 2 (UAAM); Franklin Co., Shores Lake, Ozark National Forest, 35.63 -93.95, 20-V-1983, J.M. Campbell, 1 (CNC); Grant Co., junction of highway 167 and 35 (2.7 mi SE on 35), 34.23 -92.37, pine, berlese, 77-06, 19-II-1977, Chenowith, 1 (UAAM); Hempstead Co., 12-III-1959, 1 (FMNH); 12-III-1959, leaf trash, GO.848, 1 (FMNH); 12-III-1959, leaf trash, GO.942, 1 (FMNH); Johnson Co., 25.3 mi W of junction of highways 21 and 1003, 35.68 -93.67, pitfall trap, 73-534 Line 14, 31-V-1973, R.T. Allen, 1 (UAAM); Newton Co., hardwood forest, pitfall trapline, 75-0149, 16-VII-1975, 1 (UAAM); pitfall, 75-0159, 22-VII-1975, 1 (UAAM); 2.7 mi W of Buffalo River at highway 74, 36.00 -93.38, pitfall trap, Line 2, 12-VII-1973, D. Carlyle, 1 (UAAM); Pope Co., Pope and Van Buren Co. Lines, 12.4 mi W on highway 16, 35.70 -92.92, pitfall trap, Line 10, 21-VI-1973, 1 (UAAM); 17.3 mi W of intersection of Pope and Van Buren Co. Lines, 35.70 -92.92, pitfall trap, Line 11, 21-VII-1973, 2 (UAAM); 17.3 mi W of intersection of Pope and Van Buren Co. Lines, 35.70 -92.92, pitfall trap, 73-651 Line 11, 28-VI-1973, R.T. Allen, 1 (UAAM); Pulaski Co., Little Rock (Marsh Rd.), 34.72 -92.48, berlese leaf litter, 14-II-2000, B. Baldwin, 1 (Baldwin); Little Rock (Marsh Rd.), 34.72 -92.48, hanging sugar traps, 16-IV-2003, Baldwin, 1 (Baldwin); Union Co., pitfall trap, 75-0222 Line 3, 28-VII-1975, 1 (UAAM); 0.5 mi SE of Lapile, 33.07 -92.27, pitfall trap, 74-0136 Line 3, 10-VI-1974, Heiss, Howard and Chenowith, 1 (UAAM); 4 mi S of Mount Holly, 33.23 -92.97, pitfall trap, 74-0106 Line 15, 4-VI-1974, Heiss, Howard and Chenowith, 1 (UAAM); Washington Co., leaf trash, GO.96, 5-XII-1958, 1 (UAAM); Devils Den State Park, 35.78 -94.23, in crevice, 8-VI-1946, M.W. Sanderson, 1 (INHS); Farmington, 36.05 -94.25, black light trap, 73-46, 22-VI-1973, W.D. Wylie, 1 (UAAM); Fayetteville, N slope of Markham Hill, 36.07 -94.15, under boulder, 76-97, 4-XII-1976, R. Chenowith, 1 (UAAM). DC: Washington, Rock Creek Park, 38.97 -77.05, oak hillsides, gulley floor/log, 67-2a, 24-I-1967, W. Suter, 1 (FMNH). DE: Sussex Co., Angola, 38.68 -75.18, 20-VI-1970, 1 (AMNH). FL: Alachua Co., 4 mi SW Gainesville, Hogtown Creek, 29.65 -82.38, pitfall, 27-V-1983, R.M. Reeves, 3 (DENH); Gadsden Co., 1 mi E of Havana, 30.62 -84.41, pine-hardwood litter, 6-I-1977, C.W. O’Brien and G.B. Marshall, 2 (FMNH); Hamilton Co., 8 mi S of Jasper, Hwy 129, 30.40 -82.93, mixed hardwood litter, 24-III-1977, C.W. O’Brien et al., 1 (FMNH); Leon Co., Tallahassee, 30.43 -84.28, mixed hardwood litter, 8-IV-1976, C.W. O’Brien and G.B. Marshall, 1 (FMNH); Tallahassee, 30.43 -84.28, mixed hardwood litter, 76-115, 28-VI-1976, Marshall and Justice, 1 (UAAM); Tallahassee, 30.43 -84.28, hardwood litter, 4-III-1976, G.B. Marshall, 1 (FMNH); Tallahassee, 30.43 -84.28, hardwood litter, blacklight trap, 6-XII-1976, G.B. Marshall, 1 (FMNH); Tallahassee, 30.43 84.28, mixed hardwood litter, 20-IV-1976, C.W. O’Brien and Justice, 1 (FMNH); Tallahassee, 30.43 -84.28, mixed hardwood litter, 2-IV-1976, C.W. O’Brien and G.B. Marshall, 1 (FMNH); Tallahassee, 30.43 -84.28, hardwoods, berlesate hardwoods, 77-204, 14-X-1977, L. Justice and E. Healy, 1 (UAAM); Tallahassee, 30.43 -84.28, mixed hardwood litter, 11-III-1977, G.B. Marshall, 1 (FMNH); Tallahassee, 30.43 -84.28, mixed hardwood litter, 6-X-1977, C.W. O’Brien, 1 (FMNH); Okaloosa Co., Destin, 30.40 -86.50, 13-II-1976, E.J. Kiteley, 1 (CNC); Destin, 30.40 -86.50, 10-III-1980, E.J. Kiteley, 1 (CNC); Destin, 30.40 -86.50, 23-III-1982, E.J. Kiteley, 1 (CNC); Orange Co., Winter Park, 28.60 -81.33, 4-III-1939, F.E. Lutz, 1 (AMNH); Polk Co., in steiner trap, 14-III-1961, R.E. Vild, 1 (FSCA); Suwannee Co., 15 mi SW of Live Oak, Hwy 51, pine and hardwood litter, 24-III-1977, C.W. O’Brien et al., 1 (FMNH); Taylor Co., 5 mi E of Gainesville, Hatchet Creek, 29.65 -82.25, UV light, 27-V-1983, R.M. Reeves, 2 (DENH). GA: Fulton Co., Atlanta, 33.75 -84.38, 1-VI-1929, P.W. Fattig, 1 (INHS); Liberty Co., Riceboro, 31.73 -81.43, 27-IV-1937, 1 (CAS). IL: Champaign Co., Mahomet, Hart Memorial Woods, 40.20 -88.40, trap 4, 4-VI-1960, 1 (INHS); Mahomet, Hart Memorial Woods, 40.20 -88.40, trap 13, 1-VII-1966, 1 (INHS); Mahomet, Hart Memorial Woods, 40.20 -88.40, trap 3, 7-VI-1967, R.T. Allen and J.D. Unzicker, 1 (INHS); Clark Co., Rocky Branch, 39.47 -87.77, pitfall trap, 1 to 13-V-1996, M.A. Goodrich, 1 (EIU); Rocky Branch, 39.47 -87.77, pitfall trap, 12 to 26-VII-1998, M.A. Goodrich, 1 (EIU); Rocky Branch, 39.47 -87.77, pitfall trap, 14 to 28-VI-1998, M.A. Goodrich, 3 (EIU); Rocky Branch, 39.47 -87.77, pitfall trap, 24 to 31-V-1998, M.A. Goodrich, 1 (EIU); Coles Co., Charleston vicinity, 39.50 -88.18, 6-VI-2001, N.L. Owens, 1 (EIU); Cook Co., Des Plaines, Carlé Woods, 42.03 -87.88, rotten oak log, 12-X-1957, O. Park, 1 (FMNH); Edgar Co., 4 mi SSE of Kansas, 39.50 -87.92, ‘from part of forest NOT burned in fall 1999’, pitfall trap, 25-VIII to 1-IX-2000, M.A. Goodrich, 1 (EIU); 4 mi SSE of Kansas, 39.50 -87.92, ‘from part of forest burned in 1999’, pitfall trap, 13 to 20-X-2000, M.A. Goodrich, 1 (EIU); 4 mi SSE of Kansas, 39.50 -87.92, ‘from part of forest NOT burned in 1999’, pitfall, 31-V to 7-VI-2000, M.A. Goodrich, 1 (EIU); Jackson Co., Carbondale, 37.73 -89.22, 5-V-1959, M. Stibitz, 1 (SIUC); Carbondale, 37.73 -89.22, 28-V-1963, C.S. Adams, 1 (SIUC); Piatt Co., White Health, 40.08 -88.52, ground no. 24, 1-I-1983, J.C. Dirks, 1 (INHS); Pope Co., Dixon Springs Park, Ghost Dance Canyon Trail, 37.38 -88.67, 125m, mixed hardwood forest in ravine near stream, under rock, 1043, 22-VI-2002, A. Newton, 1 (FMNH); St. Clair Co., 4-VII-1901, G.W. Block, 1 (UMRM); 5-VI-1902, G.W. Block, 1 (UMRM); Vermillion Co., Catlin, Camp Drake, 40.10 -87.75, ground, 12-VI-1940, J.E. Porter, 1 (INHS). IN: Jasper Co., Jasper-Pulaski State Game Area, 41.15 -86.93, sweeping vegetation, grassy area, 19-V-1996, R.M. Brattain, 1 (Brattain); Tippecanoe Co., 17-V-1959, N.M. Downie, 1 (FMNH); Vigo Co., 8-VI-1893, W.S. Blatchley, 1 (PURC). KS: Douglas Co., Lecompton, 39.05 -95.40, 10-X-1933, 1 (FMNH); Jefferson Co., University of Kansas Natural History Reserve, 39.05 -95.18, forest, litter, 6-V-1986, J. Pakaluk, 1 (KSEM); Leavenworth Co., 18-V-1956, G.W. Byers, 1 (KSEM). KY: Meade Co., 0.5 mi S of Route 1158 on Route 333, 7-VI-1986, R. A. Mattingly, 1 (ULKY). LA: West Feliciana Co., Feliciana Preserve near Freeland, 30.78 -91.25, pitfall traps, 17-V-1995, D. Pashley, 4 (LSAM); Feliciana Preserve near Freeland, 30.78 -91.25, pitfall traps, 5-X-1995, D. Pashley, 4 (LSAM); Feliciana Preserve near Freeland, 30.78 -91.25, pitfall traps, 5-IX-1995, D. Pashley, 3 (LSAM); Feliciana Preserve near Freeland, 30.78 -91.25, 16-V-1995, D. Pashley, 1 (LSAM); Feliciana Preserve near Freeland, 30.78 -91.25, pitfall traps, 23-V-1995, D. Pashley, 1 (LSAM); Feliciana Preserve near Freeland, 30.78 -91.25, pitfall traps, 24-V-1995, D. Pashley, 3 (LSAM); Feliciana Preserve near Freeland, 30.78 -91.25, pitfall traps, 6-I-1995, D. Pashley, 1 (LSAM); Feliciana Preserve near Freeland, 30.78 -91.25, pitfall traps, 5-II-1995, D. Pashley, 1 (LSAM); Feliciana Preserve near Freeland, 30.78 -91.25, 6-II-1995, D. Pashley, 7 (LSAM); Kakowi 2 (T35.R1W, S72), pitfall trap, 9-VIII-1995, D. Pashley, 1 (LSAM); Tunica Hills WMA1, 30.92 -91.49, pitfall trap, 23-V-1995, D. Pashley, 1 (LSAM); Tunica Hills WMA1, 30.92 -91.49, pitfall trap, 6-II-1995, D. Pashley, 3 (LSAM); Tunica Hills WMA1, 30.92 -91.49, pitfall trap, 18-V-1995, D. Pashley, 6 (LSAM); Tunica Hills WMA1, 30.92 -91.49, pitfall trap, 17-V-1995, D. Pashley, 4 (LSAM); Tunica Hills WMA1, 30.92 -91.49, pitfall trap, 5-XI-1995, D. Pashley, 1 (LSAM); Tunica Hills WMA1, 30.92 -91.49, pitfall trap, 25-VIII-1995, D. Pashley, 1 (LSAM); Tunica Hills WMA1, 30.92 -91.49, pitfall trap, 15-VII-1995, D. Pashley, 2 (LSAM); Tunica Hills WMA 2, 30.92 -91.49, 5-XI-1995, D. Pashley, 1 (LSAM). MI: Barry Co., Barry Game Area (T3N, R10Q, Sec. 26), 42.62 -85.47, barrier pitfall, 8-V-1980, 1 (DENH); Berrien Co., Lakeside, Warren Woods, 41.83 -86.63, 6-X-1973, W. Suter, 1 (FMNH); Warren Woods State Park, 41.83 -86.63, virgin beech-maple forest, berlese, leaf and log litter, 892, 30-V-1992, A. Newton and M. Thayer, 1 (FMNH). MO: Boone Co., Ashland, 38.77 -92.25, pitfall traps, 16 to 26-VIII-1984, N. Stebbins, 1 (CMNH); Columbia, 38.95 -92.33, 9-1999, students of general entomology, (LSAM); Iron Co., Annapolis, 37.37 -90.70, forest debris, 1-IV-1956, G. Ulrich, 1 (INHS); Reynolds Co., 10 mi SW of Graniteville, Johnson Shut-ins State Park, 37.53 -90.85, in dead leaves, 22-IV-1956, J. Kingsolver, 1 (INHS); St. Louis Co., Ranken, 38.53 -90.52, 1-V-1983, E. P. Meiners, 1 (CAS); Ranken, 38.53 -90.52, 6-VII-1941, E. P. Meiners, 1 (UMRM); St. Louis, 38.63 -90.20, 29-IV-1906, G.W. Bock, 1 (UMRM); Wayne Co., Sam A Baker State Park, 37.25 -90.50, 11 to 12-V-1951, 1 (EIU). MS: Benton Co., Ashland, 34.83 -89.18, 8-IV-1977, S.C. Elliot, 1 (UMIC); Covington Co., ground trash, 18-XII-1973, J.R. McCoy, 1 (MEM); George Co., Lucedale, 30.93 -88.58, 26-V-1930, H. Deitrich, 1 (CUIC); Leakesville, 31.15 -88.58, 30-IX-1931, H. Deitrich, 1 (CUIC); Grenada Co., Grenada, 33.77 -89.82, trash samples peripheral to cultivated cotton, 17-IV-1981, R. Seward, 1 (MEM); Lafayette Co., 6 mi E of Oxford, 34.37 -89.42, 25-II-1977, M.T. McCraine, 1 (UMIC); Marion Co., ground trash, 30-XI-1973, J.R. McCoy, 2 (MEM); Oktibbeha Co., 5 mi W of Starkville, Adaton, 33.48 -88.92, oak buttress basal tree hole, 4-I-1982, W. Suter, 1 (FMNH); Perry Co., Richton, 31.35 -88.93, 23-IV-1930, H. Deitrich, 1 (CUIC); Pontotoc Co., 1 mi SE of Ecru, 34.33 -89.00, deciduous woods, pitfall trap, 18-VI-1980, W.H.Cross, 1 (MEM); Wayne Co., 2.5 mi SSE of Buckatunna, 31.5 -88.52, pit trap, 21-V-1983, P.K. Lago, 1 (UMIC). NE: Otoe Co., Nebraska City, 29-IV-1916, R.W. Dawson, 1 (UNSM). NJ: Atlantic Co., Atlantic City, 39.37 -74.42, 15-VI, 1 (MCZ); Burlington Co., Centerton, E along Rancocas Creek, 40.00 -74.87, mixed hardwood forest, dung trap (human), 2 to 16-VI-1969, A. Newton, 1 (FMNH); Rancocas State Park, 40.00 -74.85, mixed hardwood forest, litter, berlese, 28-VIII-1976, A. Newton and M. Thayer, 1 (FMNH); Cape May Co., Anglesea, 39.02 -74.80, 30-V-1930, 1 (MCZ). NY: Suffolk Co., Long Island, Bellport, 40.75 -72.93, 8-VI-1913, A. Nicolay, 1 (AMNH); Long Island, Sag Harbor, 41.00 -72.30, 2-V-1929, R.L., 1 (CUIC); Long Island, Wildwood State Park, 40.97 -72.80, 7-VI-1951, H. Dietrich, 1 (CUIC); Long Island, Wyandanch, 40.75 -73.37, 10-X-1915, F. M. Schott, 1 (AMNH). OH: Preble Co., Hueston Woods State Park, 39.58 -82.77, litter, beech-maple forest, 19-V-1990, P. Kovarik, 1 (FMNH). OK: Latimer Co., V-1987, K. H. Stephan, 1 (Stephan); V-1995, K. H. Stephan, 1 (Stephan); S of Red Oak, 34.95 -95.08, IV-1997, K.H. Stephan, 1 (TAMU). PA: Bucks Co., NE of Jamison, Neshaminy Creek, Horseshoe Bend, 40.27 -75.08, 1-IV-1955, W. Ivie, 1 (AMNH); Delaware Co., Chester, 39.85 -75.35, IV-1904, 1 (CUIC); Philadelphia Co., Chestnut Hill, 40.07 -75.20, 6-VII, 1 (MCZ); Chestnut Hill, 40.07 -75.20, 24-VI, 1 (MCZ). SC: Florence Co., Florence, 34.2 -79.77, 10-I-1938, 1 (CAS); Florence, 34.2 -79.77, 46m, 25-II-1938, 1 (MCZ); Florence, 34.2 -79.77, 31m, 14-II-1938, C.W.P., 1 (MCZ); Florence, 34.2 -79.77, woods trash, 16-V-1963, V.M. Kirk, 1 (FMNH); Horry Co., 3.5 mi NW of Longs, Nelson Property, 33.97 -78.77, wet mixed conifer-hardwood forest, ex. leaf litter, 1-4pm, 19-III-2000, A.S. Ramsdale, 1 (IRCW). TX: Angelina Co., 8 mi W of Lufkin, 31.33 -94.83, 21 to 25-IV-1976, A. Smetana, 1 (CNC); 8 mi W of Lufkin, 31.33 -94.83, 21-IV-1976, A. Smetana, 1 (CNC); Jasper Co., 3 mi N of Buna, US 96, 30.43 -93.97, 6-XII-1968, G.E. Ball, 1 (UASM); San Jacinto Co., Big Creek Scenic Area, 30.52 -95.10, rotting log, 18-II-1955, D.K. Rudd, 1 (TAMU); Trinity Co., 2.5 mi NW of Apple Springs, 31.22 -94.97, 23-IV-1976, A. Smetana, 2 (CNC); Tyler Co., 2.8 mi W of Spurger, Big Thicket Nat. Pres., Beech Woods Trail, 30.70 -94.18, beech-magnolia forest, berlese leaf litter, 8-III-1989, R. Anderson, 2 (FMNH). VA: Henrico Co., 1 mi W of Elko, Elko Nat. Area, 37.47 -77.23, 15-VI-1990, C.A. Pague, 1 (VMNH); Mecklenburg Co., Elm Hill S.G.M.A., DF site near Kerr Dam, 36.60 -78.30, 5 to 19-VI-1991, VMNH survey, 2 (VMNH); Prince William Co., Occoquan, 38.68 -77.27, 2-V-1943, E.S. Ross, 1 (CNC); Virginia Beach Co., Little Creek Amphibious Base, 36.92 -76.17, 12-V-1989, K.A. Buhlmann, VDNH survey, 1 (VNMH); Virginia Beach, Seashore State Park, 36.90 -76.02, scrub pitfall site, 21-VI-1989, K.A. Buhlmann VDNH survey, 5 (VNMH).
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