Canadian Journal of Arthropod Identification

The Cryptophagidae of Canada and the northern United States of America

CJAI 40 -- December 20, 2019

Georges Pelletier & Christian Hébert

An Editorial Corrigendum has been published for this paper doi:10.3752/cjai.2019.40ed.

| Abstract | Introduction | Materials & Methods | Checklist | DNA Barcoding | Taxonomy | Key to Species | Acknowledgments | References | PDF | Cite |
| Supplemental Material |

We thank the following persons for their contribution to the present publication:
Colin Johnson, world specialist who has been working on the family since the 1960s, for providing to us a useful key to identify most European species of Atomaria. This key was tremendously helpful for building the Nearctic key and identifying the main characters that can help separate each species. He provided us with an updated unpublished checklist of Palearctic species, which also included Holarctic species. He put us in contact with the Manchester Museum, where he has worked most of his life.
Richard Leschen, from the New Zealand Arthropods Collection at Landcare Research, for kindly revising this manuscript. He collaborated closely with Colin Johnson and is the main contributor to the unpublished world checklist of Coleoptera. Both men helped build the foundation of the present work.
Floyd Shockley, from the National Museum of Natural History of the Smithsonian Institution, Washington, DC, for providing holotypes and paratypes of 82 species of Cryptophagidae, many being synonymized.
Lech Borowiec, from the University of Wroclaw’s Insect Collection, Poland, for providing images of 55 species of Palearctic Cryptophagidae, 20 of them serving as illustrations in this publication.
Dmitri Logunov, curator of the Manchester Museum in England, for sending specimens of 50 Palearctic species for comparison with Nearctic specimens.
Yves Bousquet and Patrice Bouchard, from Agriculture and Agri-Food Canada’s Canadian National Collection of Insects, Arachnids and Nematodes, Ottawa, ON, for providing 5887 specimens.
Claude Chantal, private beetle collector from Varenne, Québec, for providing 1424 specimens.
Colin Favret and Étienne Normandin, from the Collection entomologique Ouellet-Robert of the Centre sur la biodiversité de l’Université de Montréal, Québec, for providing 577 specimens.
Joseph V. McHugh and Richard Hoebeke, from the University of Georgia Collection of Arthropods, Athens, Georgia, for providing 517 specimens.
Derek Sikes, from the University of Alaska Museum Insect Collection, Fairbanks, Alaska, for providing 486 specimens.
Stéphanie Boucher, from the Lyman Entomological Museum, McGill University, Ste-Anne-de-Bellevue, Québec, for providing 474 specimens.
Mario Fréchette and Marie-Chantal Émond, from the Collection d’insectes du Québec, Ministère des Forêts, de la Faune et des Parcs, Québec, Quebec, for providing 350 specimens.
Dave Langor and James Hammond, from the Northern Forestry Centre, Canadian Forest Service, Edmonton, Alberta, for providing 337 specimens.
Karen Needham, from the University of British Columbia’s Spencer Entomological Collection, Vancouver, British Columbia, for providing 252 specimens.
Reggie Webster, collector, from the Canadian Forest Service, Atlantic Forestry Centre, Fredericton, New Brunswick, for providing 126 specimens.
Steve Marshall and S.M. Paiero, from the University of Guelph, Ontario, for providing 106 specimens and one photograph.
Audrey Malachowsky, from the Staten Island Museum, New York, for providing holotypes of two species of Atomaria.
Crystal Maier, from the Field Museum of Natural History, Chicago, Illinois, for providing the paratype of Glyptophorus mycetoecus.
Jaakko Mattila, from the Finnish Museum of Natural History, Helsinki, Finland, for providing specimens of five species of northern Europe, some known to occur in North America.
Jean-Marie Perron, from the Collection d’insectes de l’Université Laval, Québec, Quebec, for providing 96 specimens, mostly from the Léon-Provancher Historical Collection, dating from the 19th century.
Allison Brown, Sean Prosser and Tyler Elliott, from the BOLD team of University of Guelph, Ontario, for producing the DNA barcode of many species of Cryptophagidae, and Megan Milton, also from the BOLD team, for her help with BOLD tools to analyze the genetic sequences of many species.
Philippe Villemaire and André Beaudoin, from the Laurentian Forestry Centre, Canadian Forest Service, Québec, Quebec, for their help with the software ArcGIS, Don Stewart, who helped us prepare the material for the DNA analysis and Isabelle Lamarre, from the same Centre, for kindly revising the introduction of the manuscript.
Frank and Waltraud Koehler, from the Koleopterologie site which has very nice photographs of live Palearctic beetles. We have used photos, with permission of the photographers, from this website: F. Koehler (3), I. Altmann (5), A. de Wilde (1), and J. Dvorak (1). (
Smithsonian Institution, for providing the photographic system, including a high-quality digital camera for taking photos of specimens.
Heather Proctor, Editor-in-Chief of the Canadian Journal of Arthropod Identification, Morgan Jackson and Robert Anderson, responsible for the Coleoptera manuscript revisions.