Morphological Summary

females
Body. 
2.9-3.4(-3.8) mm long, 2.4-2.7 times longer than wide; pronotum 1.0-1.2 times longer than wide.
Head. 
Epistomal margin with uniseriate row of tubercles with elongate mesal tubercle or with gap at midline. Frons outline convex in lateral view; vestiture fine (not hiding part of integument); surface sculpture near epistoma densely tuberculate-punctate or with isolated tubercles; central carina present; central tubercle absent or present and single, separated from base of epistomal setae by 0.5-3 tubercle diameters, without pair of circular tubercles on either side of midline; transverse carina absent; frons central fovea present or absent; circular tubercles above top of eyes present – up to, or more than one third of all tubercles. Vertex and pronotum with stridulatory apparatus  (pars stridens). Antennal club sutures bisinuate.
Prothorax. 
Protibiae with three socketed teeth on apical half (does not include apical spine).
Elytra. Interstriae impunctate, rarely punctate (observed on interstriae 2 and 3 on middle third of elytral disc), punctures 0.3-0.5 times diameter of adjacent strial punctures (punctures and striae measured at steepest part of puncture wall), interstrial setae shorter than width of scutellar shield, interstriae 3-5 times as wide as adjacent striae. Elytral declivity with four spines per side, spine 2 or 3 largest; spine 1 (largest on 2nd interstria) closer to suture than spine 2; spines 1 and 2 separated at base by distance greater than height of spine 1; spine 2 closer to spine 3 than spine 1; spine 3 tapered, apex acute or right-angled, with apical half symmetrical in lateral view; spines 2 and 3 on or not on shared tumescence, in line with spines 1 and 4 (posterodorsal view); declivital integument shiny.

males
Body. 2.9-3.4(-3.8) mm long, 2.4-2.7 times longer than wide; pronotum 1.0-1.2 times longer than wide.
Head. Epistomal margin with uniseriate row of tubercles with elongate mesal tubercle. Frons outline convex in lateral view; vestiture fine (not hiding part of integument); surface sculpture near epistoma densely tuberculate-punctate or with isolated tubercles; central carina present; central tubercle absent or present and single, separated from base of epistomal setae by 2-4(-5) tubercle diameters, without pair of circular tubercles on either side of midline; transverse carina absent; frons central fovea absent; circular tubercles above top of eyes present – up to, or more than one third of all tubercles. Vertex and pronotum without stridulatory apparatus  (pars stridens). Antennal club sutures bisinuate.
Prothorax. Protibiae with three socketed teeth on apical half (does not include apical spine).
Elytra. Interstriae impunctate, rarely punctate (observed on interstriae 2 and 3 on middle third of elytral disc), punctures 0.3-0.5 times diameter of adjacent strial punctures (punctures and striae measured at steepest part of puncture wall), interstrial setae shorter than width of scutellar shield, interstriae 3-5 times as wide as adjacent striae. Elytral declivity with four spines per side, spine 3 largest; spine 1 (largest on 2nd interstria) closer to suture than spine 2; spines 1 and 2 separated at base by distance greater than height of spine 1; spine 2 closer to spine 3 than spine 1; spine 3 pedunculate (capitate), apex acute or right-angled, with apical half asymmetrical in lateral view; spines 2 and 3 on or not on shared tumescence, in line with spines 1 and 4 (posterodorsal view); declivital integument shiny.

Geographic Distribution

Guatemala; Honduras; Mexico (Chiapas, Chihuahua, Distrito Federal, Durango, Hidalgo, Jalisco, Mexico, Michoacan, Morelos, Nuevo Leon, Oaxaca, Puebla, Tamaulipas, Tlaxcala, Veracruz, Zacatecas); USA (Arizona)

Hosts

Pinus spp.

References

Cognato, A.I.2015. Biology, systematics, and evolution of IpsIn Bark beetles: biology and ecology of native and invasive species. Edited by F.E. Vega and R.W. Hofstetter. Elsevier, San Diego, California. Pp. 351–370.

Wood, S.L. 1982. The bark and ambrosia beetles of North and Central America (Coleoptera: Scolytidae), a taxonomic monograph. Great Basin Naturalist Memoirs6: 1–1359.

Internet resources

https://www.barkbeetles.info/regional_chklist_target_species.php?lookUp=1699